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Konrad Lorenz Institut für Vergleichende Verhaltensforschung

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** = Publikationen gelistet in SCI/SSCI/Pubmed


** Batary, P; Winkler, H; Baldi, A (2004): Experiments with artificial nests on predation in reed habitats. J Ornithol (145), 1 59-63.

** Carroll, LS; Meagher, S; Morrison, L; Penn, DJ; Potts, WK (2004): Fitness effects of a selfish gene (the Mus t complex) are revealed in an ecological context. Evolution. 2004; 58(6):1318-1328

** Danchin, E; Giraldeau, LA; Valone, TJ; Wagner, RH (2004): Public information: from nosy neighbors to cultural evolution. Science. 2004; 305(5683):487-491

** Helfenstein, F; Danchin, E; Wagner, RH (2004): Assortative mating and sexual size dimorphism in Black-legged Kittiwakes. Waterbirds (27), 3 350-354.

** Helfenstein, F; Danchin, E; Wagner, RH (2004): Is male unpredictability a paternity assurance strategy. Behaviour (141) 675-690.

** Helfenstein, F; Tirard, C; Danchin, E; Wagner, RH (2004): Low frequency of extra-pair paternity and high frequency of adoption in Black-legged Kittiwakes. Condor (106), 1 149-155.

** Hoi, H; Kristin, A; Valera, F; Hoi, C (2004): Clutch enlargement in lesser gray shrikes (Lanius minor) in Slovakia when food is superabundant: A maladaptive response. Auk (121), 2 557-564.

** Kruckenhauser, L; Haring, E; Pinsker, W; Riesing, MJ; Winkler, H; Wink, M; Gamauf, A (2004): Genetic vs. morphological differentiation of Old World buzzards (genus Buteo, Accipitridae). Zool Scr (33), 3 197-211.

** Schmidt, V; Schaefer, HM; Winkler, H (2004): Conspicuousness, not colour as foraging cue in plant-animal signalling. Oikos (106), 3 551-557.

** Tebbich, S; Taborsky, M; Fessl, B; Dvorak, M; Winkler, H (2004): Feeding behavior of four arboreal Darwin's finches: Adaptations to spatial and seasonal variability. Condor (106), 1 95-105.

** Valera, F; Hoi, H; Darolová, A; Kristofik, J (2004): Size versus health as a cue for host choice: a test of the tasty chick hypothesis. Parasitology. 2004; 129(Pt 1):59-68

** Vogl, W; Taborsky, B; Taborsky, M; Teuschl, Y; Honza, M (2004): Habitat and space use of European cuckoo females during the egg laying period. Behaviour (141) 881-898.

** Wagner, RH; Helfenstein, F; Danchin, E (2004): Female choice of young sperm in a genetically monogamous bird. Proc Biol Sci. 2004; 271 Suppl 4:S134-S137
[Open Access]

** Winkler, H; Leisler, B; Bernroider, G (2004): Ecological constraints on the evolution of avian brains. J Ornithol (145), 3 238-244.

** Zala, SM; Penn, DJ (2004): Abnormal behaviours induced by chemical pollution: a review of the evidence and new challenges. Anim Behav (68) 649-664.

** Zala, SM; Potts, WK; Penn, DJ (2004): Scent-marking displays provide honest signals of health and infection. Behav Ecol (15), 2 338-344.

** Zampiga, E; Hoi, H; Pilastro, A (2004): Preening, plumage reflectance and female choice in budgerigars. Ethol Ecol Evol (16), 4 339-349.

Publizierter (zitierfähiger) Beitrag für wissenschaftliche Veranstaltung in Proceedings (A2)

Zala, SM (2004): Mate choice, genetic resistance to infectious diseases, and modulation of testosterone. -10th International Behavioral Ecology Congress; 2004; Jyväskylä, Finland .

Originalbeitrag in Sammelwerk

Winkler, H; Leisler, B; Greenberg, R; Marra, PP (2004): To be a migrant: ecomorphological burdens and chances. IN: Greenberg, R [Hrsg.]: Birds of Two Worlds: The Ecology and Evolution of Migratory Birds.. Baltimore, Princeton University Press, pp. 79-86. ISBN: 0-8018-8107-2.


** Heindl, M; Winkler, H (2003): Vertical lek placement of forest-dwelling manakin species (Aves, Pipridae) is associated with vertical gradients of ambient light. Biol J Linn Soc (80), 4 647-658.

** Heindl, M; Winkler, H (2003): Interacting effects of ambient light and plumage color patterns in displaying Wire-tailed Manakins (Aves, Pipridae). Behav Ecol Sociobiol (53), 3 153-162.

** Heindl, M; Winkler, H (2003): Female canaries (Serinus canaria) associate more with males that contrast strongly against the background. Ethology (109), 3 259-271.

** Helfenstein, F; Wagner, RH; Danchin, E (2003): Sexual conflict over sperm ejection in monogamous pairs of kittiwakes Rissa tridactyla. Behav Ecol Sociobiol (54), 4 370-376.

** Helfenstein, F; Wagner, RH; Danchin, E; Rossi, JM (2003): Functions of courtship feeding in black-legged kittiwakes: natural and sexual selection. Anim Behav (65) 1027-1033.

** Hirschenhauser, K; Winkler, H; Oliveira, RF (2003): Comparative analysis of male androgen responsiveness to social environment in birds: the effects of mating system and paternal incubation. Horm Behav. 2003; 43(4):508-519

** Hoi, H; Vaclav, R; Slobodova, D (2003): Postmating sexual selection in house sparrows: can females estimate "good fathers" according to their early paternal effort. Folia Zool (52), 3 299-308.

** Honza, M; Taborsky, B; Taborsky, M; Teuschl, Y; Vogl, W; Moksnes, A; Roskaft, E (2003): Behaviour of female common cuckoos, Cuculus canorus, in the vicinity of host nests before and during egg laying: a radiotelemetry study. (vol 64, pg 861, 2002). Anim Behav (65) 1057-1057.

** Penn, DJ (2003): The evolutionary roots of our environmental problems: toward a Darwinian ecology. Q Rev Biol. 2003; 78(3):275-301

** Schaefer, HM; Schmidt, V; Winkler, H (2003): Testing the defence trade-off hypothesis: how contents of nutrients and secondary compounds affect fruit removal. Oikos (102), 2 318-328.

** Vaclav, R; Hoi, H; Blomqvist, D (2003): Food supplementation affects extrapair paternity in house sparrows (Passer domesticus). Behav Ecol (14), 5 730-735.

** Valera, F; Casas-Crivillé, A; Hoi, H (2003): Interspecific parasite exchange in a mixed colony of birds. J Parasitol. 2003; 89(2):245-250

** Valera, F; Hoi, H; Kristin, A (2003): Male shrikes punish unfaithful females. Behav Ecol (14), 3 403-408.

** Wagner, RH; Danchin, E (2003): Conspecific copying: a general mechanism of social aggregation. Anim Behav (65) 405-408.

Publizierter (zitierfähiger) Beitrag für wissenschaftliche Veranstaltung in Proceedings (A2)

Penn, DJ; Zala, SM (2003): Pathological alterations in behavior due to industrial pollution: Another aspect of global change. -ASAB Summer-Meeting; 2003; Grünau, Austria.

Zala, SM (2003): Testing the immunocompetence handicap hypothesis. -ASAB Summer-Meeting; 2003; Grünau, Austria.

Originalbeitrag in Sammelwerk

** Leisler, B; Winkler, H (2003): Morphological consequences of migration in passerines. Symposium on Avian Migration, CONSTANCE, GERMANY, SEP 03-06, 2001. IN: Berthold, P; Gwinner, E; Sonnenschein, E [Hrsg.]: Avian Migration. Berlin, Springer, pp. 175-186. ISBN: 3-540-43408-9.

Wagner, RH (2003): Social uses of copulation in socially monogamous razorbills. IN: Reichard, U [Hrsg.]: Monogamy: mating strategies and partnerships in birds, humans and other mammals.. Cambridge, Cambridge University Press, pp. 95-108. ISBN: 0-5215-2577-2.

Wagner, RH (2003): Social uses of copulation in socially monogamous razorbills. IN: Reichard, U [Hrsg.]: Monogamy: mating strategies and partnerships in birds, humans and other mammals:. Cambridge, Cambridge University Press, pp. 95-108. ISBN: 978-0-5218-1973-2.


** Hoi, H; Hoi, C; Kristofik, J; Darolova, A (2002): Reproductive success decreases with colony size in the European bee-eater. Ethol Ecol Evol (14), 2 99-110.

** Mettke-Hofmann, C; Winkler, H; Leisler, B (2002): The significance of ecological factors for exploration and neophobia in parrots. Ethology (108), 3 249-272.

** Romero-Pujante, M; Hoi, H; Blomqvist, D; Valera, F (2002): Tail length and mutual mate choice in bearded tits (Panurus biarmicus). Ethology (108), 10 885-895.

** Schmidt, KT; Hoi, H (2002): Supplemental feeding reduces natural selection in juvenile red deer. Ecography (25) 265-272.

** Vaclav, R; Hoi, H (2002): Different reproductive tactics in house sparrows signalled by badge size: Is there a benefit to being average. Ethology (108), 7 569-582.

** Vaclav, R; Hoi, H (2002): Importance of colony size and breeding synchrony on behaviour, reproductive success and paternity in house sparrows Passer domesticus. Folia Zool (51), 1 35-48.

** Vaclav, R; Hoi, H; Blomqvist, D (2002): Badge size, paternity assurance behaviours and paternity losses in male house sparrows. J Avian Biol (33), 3 315-318.

** Vogl, W; Taborsky, M; Taborsky, B; Teuschl, Y; Honza, M (2002): Cuckoo females preferentially use specific habitats when searching for host nests. Anim Behav (64) 843-850.

Ille, R; Hoi, H; Grinschgl, F; Zink, R (2002): Paternity assurance in two species of colonially breeding falcon: the kestrel Falco tinnunculus and the red-footed falcon Falco vespertinus. Etología (10) 11-15.

Originalbeitrag in Sammelwerk

Winkler, H; Christie, D (2002): Order Piciformes; Family Picidae. IN: Hoyo, J. [Hrsg.]: Handbook of the Birds of the World. Barcelona, Lynx Edicions, pp. 296-555. ISBN: 84-87334-37-7.


** Darolova, A; Hoi, H; Kristofik, J; Hoi, C (2001): Horizontal and vertical ectoparasite transmission of three species of Malophaga, and individual variation in european bee-eaters (Merops apiaster). J Parasitol. 2001; 87(2):256-262

** Hoi-Leitner, M; Romero-Pujante, M; Hoi, H; Pavlova, A (2001): Food availability and immune capacity in serin (Serinus serinus) nestlings. Behav Ecol Sociobiol (49), 5 333-339.

** Michalek, KG; Winkler, H (2001): Parental care and parentage in monogamous great spotted woodpeckers (Picoides major) and middle spotted woodpeckers (Picoides medius). Behaviour (138) 1259-1285.

** Nemeth, E; Winkler, H; Dabelsteen, T (2001): Differential degradation of antbird songs in a neotropical rainforest: adaptation to perch height? J Acoust Soc Am. 2001; 110(6):3263-3274

** Palme, R; Entenfellner, U; Hoi, H; Möstl, E (2001): Faecal oestrogens and progesterone metabolites in mares of different breeds during the last trimester of pregnancy. Reprod Domest Anim. 2001; 36(5):273-277

** Valera, F; Kristin, A; Hoi, H (2001): Why does the lesser grey shrike (Lanius minor) seldom store food Determinants of impaling in an uncommon storing species. Behaviour (138) 1421-1436.

** Winkler, H; Kothbauer-Hellmann, R (2001): The role of search area in the detection of cryptic prey by crested tits and coal tits. Behaviour (138) 873-883.

** Winkler, H; Preleuthner, M (2001): Behaviour and ecology of birds in tropical rain forest canopies. Meeting of the European-Science-Foundation Canopy Research Programme, OXFORD, ENGLAND, ,. Plant Ecol (153), 1-2 193-202.

Buch, Monografie

Hoi, H (2001): The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 177. ISBN: 3-7001-3026-0.

Originalbeitrag in Sammelwerk

Hoi, H; Darolova, A; Kristofik, J (2001): Factors influencing nest depredation in European reed passerines. IN: Hoi, H; [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 27-43.

Hoi, H; Hoi, C (2001): Intrasexual selection in bearded tits: Beard length - a reliable predictor for dominance rank. IN: Hoi, H. [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 121-138.

Hoi, H; Hoi, C (2001): Habitat selection and habitat use of the bearded tit (Panurus biarmicus). IN: Hoi, H; [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 73-86. ISBN: 3-7001-3026-0.

Hoi, H; Ille, R (2001): The role of simple song in the marsh warbler (Acrocephalus palustris). IN: Hoi, H; [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 111-119.

Hoi, H; Ille, R; Hoi-Leitner, M.; Kleindorfer, S (2001): An experimental study of short-term parental feeding adjustment in Acrocephalus warblers. IN: Hoi, H; [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 161-177.

Kleindorfer, S; Hoi, H.; Ille, R; Hoi-Leitner, M (2001): An experimental study of short-term parental feeding adjustment in Acrocephalus warblers. IN: Hoi, H. [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 161-177. ISBN: 3-7001-3026-0.

Romero-Pujante, M; Winkler, H; Hoi, H (2001): The role of tail length for flight performance in the bearded tit (Panurus biarmicus): A tail elongation experiment. IN: Hoi, H. [Hrsg.]: The ecology of reed birds. Wien, Österreichische Akademie der Wissenschaften, pp. 151-160. ISBN: 3-7001-3026-0.

Winkler, H (2001): Der Flug der Vögel: von Lorenz bis zur modernen Ökomorphologie. IN: Kotrschal, K [Hrsg.]: Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht. Fürth, Filander, pp. 195-210. ISBN: 3-930831-33-3.

Winkler, H (2001): The ecology of avian acoustical signals. IN: Barth, FG [Hrsg.]: Ecology of sensing . Berlin, Springer, pp. 79-104. ISBN: 978-3-540-66901-2.


** Fessl, B; Hoi, H (2000): Song complexity and song structure in the Moustached Warbler Acrocephalus melanopogon. J Avian Biol (31), 2 144-150.

** Kristin, A; Hoi, H; Valera, F; Hoi, C (2000): Breeding biology and breeding success of the Lesser Grey Shrike Lanius minor in a stable and dense population. Ibis (142), 2 305-311.

** Wagner, RH; Danchin, E; Boulinier, T; Helfenstein, F (2000): Colonies as byproducts of commodity selection. Behav Ecol (11), 5 572-573.

** Hoi-Leitner, M; Hoi, H; Romero-Pujante, M; Valera, F (1999): Multi-male display sites in serins (Serinus serinus L.). Ecoscience (6), 2 143-147.

** Hoi-Leitner, M; Hoi, H; Romero-Pujante, M; Valera, F (1999): Female extra-pair behaviour and environmental quality in the serin (Serinus serinus): a test of the 'constrained female hypothesis'. P Roy Soc Lond B Bio (266), 1423 1021-1026.

** Lotem, A; Wagner, RH; Balshine-Earn, S (1999): The overlooked signaling component of nonsignaling behavior. Behav Ecol (10), 2 209-212.

** Schmidt, KT; Hoi, H (1999): Feeding tactics of low-ranking red deer stags at supplementary feeding sites. Ethology (105), 4 349-360.

Publizierter (zitierfähiger) Beitrag für wissenschaftliche Veranstaltung in Proceedings (A2)

Danchin, E; Wagner, RH (1999): Commodity selection and coloniality: exaggerated traits are produced by processes of choice. -22nd International Ornithological Congress; AUG 16-22, 1998; Durban, SOUTH AFRICA.

Wagner, RH (1999): Sexual selection and colony formation. -22nd International Ornithological Congress; AUG 16-22, 1998; Durban, SOUTH AFRICA.


** Gamauf, A; Preleuthner, M; Winkler, H (1998): Philippine birds of prey: Interrelations among habitat, morphology, and behavior. Auk (115), 3 713-726.

** Hoi, H; Darolova, A; Konig, C; Kristofik, J (1998): The relation between colony size, breeding density and ectoparasite loads of adult European bee-eaters (Merops apiaster). Ecoscience (5), 2 156-163.

** Kleindorfer, S; Hoi, H; Fessl, B (1998): Alarm calls and chick reactions: distinguishing between levels of analysis. A reply to Curio. Anim Behav (56), 1 262-264.

** Kleindorfer, S; Hoi, H; Fessl, B (1998): Clarification of the chick reaction hypothesis. Anim Behav (55), 2 504-507.

** Sax, A; Hoi, H (1998): Individual and temporal variation in cloacal protuberance size of male Bearded Tits (Panurus biarmicus). Auk (115), 4 964-969.

** Sax, A; Hoi, H; Birkhead, TR (1998): Copulation rate and sperm use by female bearded tits, Panurus biarmicus. Anim Behav (56), 5 1199-1204.

** Schmidt, KT; Seivwright, LJ; Hoi, H; Staines, BW (1998): The effect of depletion and predictability of distinct food patches on the timing of aggression in red deer stags. Ecography (21), 4 415-422.

Originalbeitrag in Sammelwerk

Wagner, RH (1998): Hidden leks: sexual selection and the clustering of avian territories. IN: Parker, P.G.; Burley, N.; [Hrsg.]: Ornithological Monograph, Vol 49.: Avian reproductive tactics: female and male perspectives.. Washington, D.C., American Ornithologists' Union, pp. 123-145.


** Darolova, A; Hoi, H; Schleicher, B (1997): The effect of ectoparasite nest load on the breeding biology of the Penduline Tit Remiz pendulinus. Ibis (139), 1 115-120.

** Hoi, H (1997): Assessment of the quality of copulation partners in the monogamous bearded tit. Anim Behav (53) 277-286.

** Hoi, H; HoiLeitner, M (1997): An alternative route to coloniality in the bearded tit: Females pursue extra-pair fertilizations. Behav Ecol (8), 2 113-119.

** Hoi, H; Ille, R (1997): Does insect sampling reflect food availability in Marsh Warblers Acrocephalus palustris a reply. Ibis (139), 4 699-701.

** Kleindorfer, S; Hoi, H (1997): Nest predation avoidance: An alternative explanation for male incubation in Acrocephalus melanopogon. Ethology (103), 8 619-631.

** Kleindorfer, S; Hoi, H; Ille, R (1997): Nestling growth patterns and antipredator responses: a comparison between four Acrocephalus warblers. Biologia (52), 5 677-685.

** KothbauerHellmann, R; Winkler, H (1997): Prey detection in two tit species, Parus ater and P-cristatus. Ethology (103), 4 339-349.

** Schleicher, B; Hoi, H; Valera, F; HoiLeitner, M (1997): The importance of different paternity guards in the polygynandrous penduline tit (Remiz pendulinus). Behaviour (134) 941-959.

** Valera, F; Hoi, H; Schleicher, B (1997): Egg burial in penduline tits, Remit pendulinus: Its role in mate desertion and female polyandry. Behav Ecol (8), 1 20-27.

Originalbeitrag in Sammelwerk

Wagner, RH (1997): Hidden Leks: Sexual Selection and the Clustering of Avian Territories. IN: Parker, P; Burley, N; [Hrsg.]: Ornithological Monographs. Washington, DC, American Ornithologists Union, pp. 123-145.


** Darolova, A; Hoi, H (1996): The importance of nest predation and food availability in penduline tits. Folia Zool (45), 3 237-246.

** Fessl, B; Hoi, H (1996): The significance of a two part song in the moustached warbler (Acrocephalus melanopogon). Ethol Ecol Evol (8), 3 265-278.

** Fessl, B; Kleindorfer, S; Hoi, H; Lorenz, K (1996): Extra male parental behaviour: Evidence for an alternative mating strategy in the Moustached Warbler Acrocephalus melanopogon. J Avian Biol (27), 1 88-91.

** Grubbauer, P; Hoi, H (1996): Female penduline tits (Remiz pendulinus) choosing high quality nests benefit by decreased incubation effort and increased hatching success. Ecoscience (3), 3 274-279.

** Hoi, H; Ille, R (1996): Trade-offs of territory choice in male and female marsh warblers. Auk (113), 1 243-246.

** Hoi, H; Schleicher, B; Valera, F (1996): Nest size variation and its importance for mate choice in penduline tits, Remit pendulinus. Anim Behav (51) 464-466.

** Kleindorfer, S; Hoi, H; Fessl, B (1996): Alarm calls and chick reactions in the moustached warbler, Acrocephalus melanopogon. Anim Behav (51) 1199-1206.

** Tebbich, S; Taborsky, M; Winkler, H (1996): Social manipulation causes cooperation in keas. Anim Behav (52) 1-10.

** Hoi, H; Kleindorfer, S; Ille, R; Dittami, J (1995): Prey Abundance And Male Parental Behavior In Acrocephalus Warblers. Ibis (137), 4 490-496.

** Hoileitner, M; Nechtelberger, H; Hoi, H (1995): Song Rate As A Signal For Nest-Site Quality In Blackcaps (Sylvia-Atricapilla). Behav Ecol Sociobiol (37), 6 399-405.

** Ille, R; Hoi, H (1995): Factors Influencing Fledgling Survival In The Marsh Warbler Acrocephalus-Palustris - Food And Vegetation Density. Ibis. 1995; 137(4): 586-588.

** Kleindorfer, S; Fessl, B; Hoi, H (1995): More Is Not Always Better - Male Incubation In 2 Acrocephalus Warblers. Behaviour (132) 607-625.

** Uiblein, F (1995): Effects Of Vertical Prey Distribution On The Food-Searching Behavior Of Danube Bleak Chalcalburnus Chalcoides Mento. 2nd International Joint Conference on Limnology and Oceanography, EVIAN, FRANCE, MAY 25-28, 1993. Hydrobiologia (300) 329-333.

** Birkhead, TR; Hoi, H (1994): Reproductive-Organs And Mating Strategies Of The Bearded Tit Panurus-Biarmicus. Ibis. 1994; 136(3): 356-359.

** Hoi, H; Schleicher, B; Valera, F (1994): Female Mate Choice And Nest Desertion In Penduline Tits, Remiz Pendulinus - The Importance Of Nest Quality. Anim Behav. 1994; 48(3): 743-746.

** Hoi, H; Winkler, H (1994): Predation on nests - a case of apparent competition. Oecologia (98), 3-4 436-440.

** Schleicher, B; Valera, F; Hoi, H (1993): The Conflict Between Nest Guarding And Mate Guarding In Penduline Tits (Remiz-Pendulinus). Ethology (95), 2 157-165.

** Winkler, H; Leisler, B (1992): On The Ecomorphology Of Migrants. CONF ON ECOLOGY AND CONSERVATION OF PALAEOARCTIC-AFRICAN MIGRANTS, NORWICK, ENGLAND, APR 04-07, 1991. Ibis (134) 21-28.

** Winkler, H; Orellana, CP (1992): The Costs Of Mixed Prey For A Planktivorous Cyprinid, Chalcalburnus-Chalcoides-Mento. Environ Biol Fish (35), 1 85-93.

** Winkler, H; Orellana, CP (1992): Functional-Responses Of 5 Cyprinid Species To Planktonic Prey. Environ Biol Fish (33), 1-2 53-62.

** Mbahinzireki, G; Uiblein, F; Winkler, H (1991): Microhabitat Selection Of Ostracods In Relation To Predation And Food. Hydrobiologia (222), 2 115-119.

** Hoi, H; Winkler, H (1988): Predation pressure on reed-breeding birds - an experimental-study. J Ornithol (129), 4 439-447.

Populärwissenschaftlicher Artikel

Hoi, H (1988): Überlegungen zum Brutpflegesystem beim Cassinschnäpper (Muscicapa cassini) - einem tropischen Fließwasserbewohner. Egretta (31) 78-82.


** Hoi, H (1987): Brood division and habitat usage in the cassins gray flycatcher (Muscicapa, Cassini). J Ornithol (128), 3 338-342.

** Catchpole, C; Leisler, B; Winkler, H (1985): Polygyny In The Great Reed Warbler, Acrocephalus-Arundinaceus - A Possible Case Of Deception. Behav Ecol Sociobiol (16), 3 285-291.

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